World-renowned chimpanzee expert Jane Goodall died of cardiopulmonary arrest, according to the death certificate obtained by TMZ.

The primatologist was 91 when she died in Los Angeles on October 1, while in the United States on a speaking tour.

Goodall’s death certificate also indicated that she had epilepsy, although it is unclear if that played a role in her death.

The Jane Goodall Institute said its founder died “due to natural causes” when it announced her death earlier this month.

In the statement, the organization said: “Dr. Goodall’s discoveries as an ethologist revolutionized science and she was a tireless advocate for the protection and restoration of our natural world.”

Jane Goodall, seen here in September, died of cardiopulmonary arrest at the age of 91 earlier this month, according to the death certificate obtained by TMZ.
Jane Goodall, seen here in September, died of cardiopulmonary arrest at the age of 91 earlier this month, according to the death certificate obtained by TMZ.

Taylor Hill via Getty Images

Goodall fell in love with chimpanzees after encountering them for the first time during a trip to Kenya when she was 22 years old. She was the first scientist to discover chimpanzees using twigs to collect termites, debunking the previous belief that only humans used tools.

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A lifelong conservationist and animal rights activist, Goodall often spoke of humans’ collective responsibility to care for the Earth, its creatures, and the people around us.

“Each of us matters, has a role to play, and makes a difference,” he wrote in his 1999 book, “Reason for Hope: A Spiritual Journey.” “Each of us must take responsibility for our own lives and, above all, show respect and love for the living beings around us, especially among ourselves.”